Updated

JACKSONVILLE, Fla. (AP) -- Jacksonville Jaguars running back T.J. Yeldon has missed another practice.

Yeldon sat out his fourth consecutive practice Wednesday because of a groin injury. The rookie missed all three workouts last week and did not play in a loss to Houston. Without him, Toby Gerhart and Denard Robinson combined for 45 yards on 16 carries.

Yeldon has 291 yards rushing, 73 yards receiving and a touchdown this season.

Receiver Marqise Lee (hamstring) and safeties Johnathan Cyprien (calf) and James Sample (shoulder) also missed practice for Jacksonville (1-5), which plays Buffalo (3-3) on Sunday in London.

Running back Bernard Pierce returned to practice after spending last week in the NFL's concussion protocol.