The only surviving witness of rapper Tupac Shakur's 1996 murder has been indicted, Fox News Digital can confirm.

Duane "Keffe D" Davis was indicted on the charge of open murder with use of a deadly weapon along with a gang enhancement, prosecutors revealed during a press conference Friday. In 2018, Davis made media statements that "reinvigorated" the investigation, police explained.

The charges were revealed Friday, hours after Davis, 60, was arrested while on a walk near his home, according to prosecutors.

Chief Deputy District Attorney Marc DiGiacomo said a grand jury had been seated in the case for "several months." The district attorney described Davis as the "on-ground, on-site commander" who "ordered the death" of Shakur.

POLICE RELEASE SWAT VIDEO OF RAID LINKED TO TUPAC SHAKUR MURDER

Duane Davis was arrested and indicted for Tupac's murder

Duane "Keffe D" Davis was indicted Friday in Tupac Shakur's 1996 murder. (LAS VEGAS METROPOLITAN POLICE DEPARTMENT)

Police also revealed Davis obtained the gun used to kill Shakur from a "close associate." Davis was described as the "shot-caller" of a group of three suspects – all now deceased – and accused of orchestrating the plan to murder the rapper.

Davis has long been known to investigators and has himself admitted in interviews and in his 2019 tell-all memoir, "Compton Street Legend," that he was in the Cadillac when the gunfire erupted during the September 1996 drive-by shooting.

WATCH: POLICE RAID HOME IN CONNECTION WITH TUPAC SHAKUR'S MURDER

Police raided the suspect's wife's Nevada home on July 17. "Come out with your hands up and your hands empty!" law enforcement officers were heard yelling in the video as blue and red lights lit up the otherwise quiet neighborhood in the city of Henderson, which is about 20 miles from the Las Vegas Strip.

Authorities seized a Pokeball USB drive, an iPhone, three iPads (one with a cracked screen), four laptops, a tablet, a desktop computer, several external hard drives, copies of the book "Compton Street Legends," a Vibe magazine about Shakur and two "black tubs" of photos, according to the search warrant.

Police also searched for "notes, writings, ledgers and other handwritten or typed documents" about anything mentioning the murder of Shakur, the documents stated.

During Friday's press conference, authorities confirmed that they seized evidence using the search warrant that "corroborated" information they had collected during the investiagtion.

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Tupac Shakur raps on stage without a shirt

Duane "Keffe D" Davis was arrested Friday morning in connection to Tupac Shakur's 1996 murder. (Raymond Boyd)

Davis' nephew, Orlando Anderson, was considered the prime suspect in Shakur's 1996 murder. Anderson denied involvement before he was killed in a separate shooting in Compton, California, in 1998.

Shakur was fatally shot in a drive-by shooting on Sept. 7, 1996. The rapper was riding shotgun in Death Row Records co-founder Marion "Suge" Knight's black BMW when a white Cadillac pulled up alongside them while they were stopped at a light. In an interview done by Davis in 2018, he claimed everyone in the Cadillac that night was a member of the South Side Compton Crips gang.

Tupac

Tupac Shakur had allegedly gotten into a brawl with a member of the South Side Compton Crips gang hours before his death. (AP Photo/Frank Wiese, File)

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Tupac Shakur back stage

Tupac Shakur was fatally shot on Sept. 7, 1996. (Raymond Boyd/Getty Images)

The gang was looking for Shakur, who had allegedly brawled with a member in the hours leading up to the fatal shooting.

Davis, who made the admission during the "Unsolved: The Murders of Tupac and the Notorious B.I.G." documentary, only revealed the gunshots came from the back seat.

Shakur was just 25 at the time. His fourth solo record, "All Eyez on Me," was still at the top of the charts with about 5 million copies sold.

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Tupac Shakur poses for a photo

Tupac Shakur remains one of the top-selling hip-hop artists decades after his death. (Al Pereira/Michael Ochs Archives/Getty Images)

Fox News Digital's Chris Eberhart and The Associated Press contributed to this report.