Updated

A man accused along with former University of Georgia football coach Jim Donnan of operating a fraudulent investment scheme testified that Donnan was aware that money from some investors was being used to pay others.

Gregory Crabtree began testifying Thursday in Donnan's trial on federal charges including conspiracy, mail fraud and wire fraud.

A federal indictment unsealed last year accused Donnan, of Athens, and Crabtree, of Proctorville, Ohio, of running the scheme through GLC Limited, Inc., a West Virginia-based company that dealt in closeout merchandise.

Crabtree pleaded guilty last month to a single charge of conspiracy.

Crabtree told the court Donnan asked him to sign a document in late 2010, after the scheme had unraveled, certifying that Donnan wasn't aware that investor money was being used to pay other investors.