Updated

A federal grand jury in Nevada has indicted Cliven Bundy and four others on 16 charges related to an armed standoff near his ranch in 2014 over unpaid grazing fees.

The 69-year-old rancher was arrested in Feb. 10 in Portland, Oregon, where his sons, Ammon and Ryan Bundy, are jailed and accused of organizing the occupation of Malheur National Wildlife Refuge. They had demanded that public lands be turned over to locals.

Ammon and Ryan Bundy, along with Ryan Payne and Peter Santilli, were also indicted Wednesday.

They and Cliven Bundy face charges including conspiracy to commit an offense against the United States and conspiracy to impede or injure a federal officer, among others.

The indictment also alleges counts of criminal forfeiture that would require the seizure of property totaling at least $3 million.

The defendants are currently in custody in Oregon.