Updated

A Congolese army spokesman says a former colonel whose fighters have been accused of raping civilians has been killed in the country's volatile east.

Col. Emmanuel Nsengiyumva was killed Saturday during a military operation in North Kivu province, Maj. Olivier Hamuli said Monday.

Nsengiyumva formerly belonged to a Tutsi-led rebel group. He later joined the Congolese army but then deserted and fought alongside various other armed groups.

A preliminary U.N. report said Nsengiyumva's fighters and other armed men "perpetrated serious human rights violations" against civilians in eastern Congo last July and August.

More than 300 people were raped in the attacks around Luvungi village, the report said.