Updated

LAS VEGAS, June 9 - Manny Pacquiao of the Philippines was stunningly upset by American Timothy Bradley on Saturday, surrendering his WBO welterweight title on a split decision at the MGM Grand Garden Arena.

It was Pacquiao's first defeat since he lost to Erik Morales in Las Vegas in March 2005, ending a run of 15 consecutive wins by the Filipino who has won world titles in an unprecedented eight weight divisions.

(Reporting by Mark Lamport-Stokes in Las Vegas; Editing by John O'Brien)