NASHVILLE, Tenn. (AP) Frank Mordica, the Vanderbilt running back who shares the single-best rushing performance in Southeastern Conference history, has died. He was 57.

His nephew told Vanderbilt officials he died Saturday of a heart attack while exercising in his hometown Tallahassee, Florida.

Mordica ran for 321 yards against Air Force on Nov. 18, 1978, setting an SEC record matched only by Darren McFadden at Arkansas against South Carolina in 2007 on 12 more carries.

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With 2,632 yards between 1976 and 1979, Mordica was Vanderbilt's career rushing leader until passed by Zac Stacy in 2012.

Drafted No. 233 overall in 1980 by the New Orleans Saints, Mordica never played because of a knee injury. After graduating, he joined the U.S. Navy before retiring in 2011.

He is survived by his son, Frank III.