Updated

State officials say a lack of oxygen caused the death of tens of thousands of fish after they likely sought shelter in a Southern California harbor.

Janice Mackey of the Department of Fish and Wildlife said Tuesday that scientists suspect the large school of anchovies may have sought cover from a predator on Saturday.

She said once in the Marina del Rey harbor, the fish depleted the water of oxygen.

Mackey says workers have removed 300 bags of fish weighing a total of more than six tons.

The dead anchovies and other fish including stingrays created a silvery blanket on the water's surface that set off a feeding frenzy among harbor seals, pelicans and seagulls.

Similar fish die-offs occurred in Ventura Harbor and at Redondo Beach in 2011.