PHILADELPHIA (AP) The Eagles have released two-time Pro Bowl linebacker DeMeco Ryans.

The move announced Wednesday saves Philadelphia $3.5 million in salary cap space. Ryans played four seasons with the Eagles after being acquired in a trade from Houston.

Ryans had 458 tackles, five sacks and five interceptions in 54 regular-season games. He was a team leader and strong presence in the locker room.

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Ryans overcame rupturing his Achilles tendon midway through the 2014 season and started 13 games last year. It was the second time he suffered that injury in his 10-year career.

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AP NFL website: www.pro32.ap.org and www.twitter.com/AP-NFL