The latest developments as Russia and other nations counter Islamic State militants in Syria. All times local:

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12:20 p.m.

A spokesman for Russian President Vladimir Putin says Russia's airstrikes in Syria are targeting not only Islamic State militants but also other terrorist groups.

Russia on Wednesday carried out its first airstrikes in Syria. Moscow insisted that it was targeting IS militants while U.S. officials and other cast doubt on the claim, saying the Russians appeared to be attacking opposition groups fighting Syrian government forces.

Dmitry Peskov told reporters on Thursday that Russia is going after IS militants and other "well-known" groups.

Asked whether Russia is choosing its own targets or following the instructions of Syrian President Bashar Assad, Peskov said the targets were chosen "in coordination" with the Syrian army.

The Russian defense ministry said late Wednesday Russia had carried out 20 airstrikes.