Backup quarterback Quinn Epperly's 36-yard touchdown pass to Roman Wilson with 13 seconds remaining lifted Princeton to a 39-34 win Saturday over Harvard, capping a 29-point fourth quarter for the Tigers and snapping the Crimson's nation-leading 14-game winning streak.
Epperly took over after Connor Michelsen, who had thrown three straight touchdown passes, left the game following a sack midway through the 90-yard drive.
The victory gave Princeton sole possession of first place in the Ivy League.
The Tigers (4-2, 3-0) overcame a career day by Harvard's Colton Chapple, who threw for a school-record 448 yards and tied his own mark of five touchdowns.
Trailing 34-10 with 13:02 left, Michelson threw a 7-yard TD to Dre Nelson, a 29-yarder to Matt Costello and a 20-yarder to Seth DeValve, closing the gap to 34-32.
Princeton won its fourth straight game and ended a five-game losing streak against the Crimson (5-1, 2-1).